Summary
It is not surprising that American vertebrate paleontology has arisen to so high a plane, when one considers the material at its disposal. Having a vast and virgin field for exploration, a sufficient number of collectors, some of whom have devoted much of their lives to the work, and a refinement of technique that permitted the preservation of the fragmental and ill conserved as well as the finer specimens, the results could hardly have been otherwise. Thus it has been possible to secure material almost unique throughout the world for extent, for completeness, and for variety. To this must be added a certain American daring in the matter of the restoration of missing portions, both of the individual bones and of the skeleton as a whole, such as European conservatism will not as a rule permit. This work has for the most part been done after the most painstaking comparison and research and is highly justified in the accuracy of the results, which render the fabric of the skeleton much more intelligible, both to the scientist and to the layman. Material once secured and prepared is then mounted, and here again American ingenuity has accomplished some remarkable results. Some of the specimens thus mounted are so small and delicate as to require holding devices comparable to those for the display of jewels; yet others—huge dinosaurs the bones of which are enormously heavy, but so brittle that they will not bear even the weight of a process unsupported—require a carefully designed and skilfully worked out series of supports of steel or iron which must be perfectly secure and at the same time as inconspicuous as possible. And of late the lifelike pose of the individual skeleton has been augmented by the preparation of groups of several animals which collectively exhibit sex, size, or other individual variations and the full mechanics of the skeleton under the varying poses assumed by the creature during life.
The work of further restoration has been rendered possible through comparative anatomical study, enabling us to essay restorations in entirety by means of models and drawings, clothing the bones with sinews and with flesh and the flesh with skin and hair, if such the creature bore; while the laws of faunal coloration have permitted the coloring of the restoration in a way which if not the actual hue of life is a very reasonable possibility.
Thus the American paleontologists have blazed a trail which has been followed to good effect by certain of their Old World colleagues.
With such means and methods and such material available, it is again not surprising that American paleontology has furnished more and more of the evidences of evolution, and disclosed to the eyes of scientists animal relationships which were undreamed of by the systematist whose research dealt only with the existing. It has also explained some vexatious problems of animal distribution and of extinction, and has connected up cause and effect in the great evolutionary movements which are recorded.
The results of systematic research have added hosts of new genera and species and of families, but of orders there are relatively few. Nevertheless a number, especially among reptiles and mammals, have come to light as the fruits of American discovery. But aside from the dry cataloguing of such groups, the American systematists have worked out some very remarkable phylogenies and have thus clarified our vision of animal relationships in a way which the recent zoologist could never have done. In this connection, the Permian vertebrates, which have been collected and studied with amazing success, principally by Williston and Case, should be mentioned, although the work is yet incomplete. Some of these forms are amphibian, others reptilian, yet others of such character as to link the two classes as transitional forms. Of the Mesozoic reptiles, a very remarkable assemblage has come to light, in a degree of perfection unknown elsewhere. These are dinosaurs, of which several phyla are now known; carnivores both great and small, some of the latter being actually toothless; Sauropoda, whose perfection and dimensions are incomparable except for those found in East Africa; and predentates, armored, unarmored, and horned, the last exclusively American. The unarmored trachodonts are now known in their entirety, for not only has our West produced articulated skeletons but mummified carcasses whose skin and other portions of their soft anatomy are represented, and which are thus far without a parallel elsewhere in the world. Other reptilian groups are well known, notably the Triassic ichthyosaurs, and the mosasaurs and plesiosaurs of the Kansas chalk. The last formation has also produced toothed birds, Hesperornis and Ichthyornis, which again are absolutely unique.
But it is in the mammalian class that the phylogenies become so highly complete and of such great importance as evolutionary evidences, for nowhere else than in our own West have such series been found as the Dinocerata and creodonts among archaic forms, the primitive primates from the Eocene, the carnivores such as the dogs and cats and mustellids, but especially the hoofed orders such as the horses. Of these hoofed orders, the classic American series of horses is complete, that of the camels probably no less so, while much is known of the deer and oreodonts, the last showing several parallel phyla, and of the proboscideans, which while having their pristine home in the Old World nevertheless soon sought the new where their remains are found from the Miocene until their final and apparently very recent extinction. These creatures show increase of bulk, perfection of feet and teeth, development of various weapons, horns and antlers, which may be studied in their relationship with the other organs to make the evolving whole, or their evolution may be traced as individual structures which have their rise, culmination, and sometimes their senile atrophy in a way comparable to that of the representatives of the order as a whole. Thus, for example, Osborn has traced the evolution of the molar teeth, and Cope of the feet, while Marsh has shown that brain development runs a similar course and that its degree of perfection within a group is a potent factor for survival.
As a student of evolution, the paleontologist sees things in a very different light from the zoologist. The latter is concerned largely with matters of detail—with the inheritance of color or of the minor and more superficial characteristics of animals—and the period of observation of such phenomena is of necessity brief because of the mortality of the observer. Whereas the paleontologist has a perspective which the other lacks, since for him time means little in the terms of his own life, and he can look into the past and see the great and fundamental changes which evolution has wrought, the rise of phyla, of classes, of orders, and he alone can see the orderliness of the process and sense the majesty of the laws which govern it.