Vertebrate Paleontology in America
Early Writers.—Having thus summarized paleontological progress in the Old World, we can turn to a consideration of the work done in the New, especially in the United States, because while the Old World investigation has been invaluable, a science of vertebrate paleontology, very complete both as to its zoological and geological scope and in the extent and value of published results, could be built exclusively upon the discoveries and researches made by Americans. The science of vertebrate paleontology may be said to have had its beginnings in North America with the activities of Thomas Jefferson, who, like Franklin, felt so strong an interest in scientific pursuits that even the graver duties of the highest office in the gift of the American people could not deter him from them. When in 1797 Jefferson came to be inaugurated as vice-president of the United States, he brought with him to Philadelphia not only his manuscript but the actual fossil bones upon which it was based. Again in 1801 he was greatly interested in the Shawangunk mastodon, despite heavy cares of state, and in 1808 made part of the executive mansion in Washington serve as a paleontological laboratory, displaying therein for study the bones of proboscideans and their contemporaries which the Big Bone Lick of Kentucky had produced. Jefferson’s work would not, perhaps, have been epoch-making were it not for its unique chronological position in the annals of the science.
Jefferson was followed by another man—this time one whose diverging lines of interest led him not into the realm of political service, but of art, for Rembrandt Peale possessed an enviable reputation among the early painters of America. Peale published in 1802 an account of the skeleton of the “mammoth,” really the mastodon, M. americanus, speaking of it as a nondescript carnivorous animal of immense size found in America. It was because of the form of the molar teeth that Peale said of it: “If this animal was indeed carnivorous, which I believe cannot be doubted, though we may as philosophers regret it, as men we cannot but thank Heaven that its whole generation is probably extinct.”
With the work of these men as a beginning, it is not strange that the more conspicuous Pleistocene fossils of the East should have attracted the attention of many subsequent writers in the first part of the nineteenth century, nor that the early papers to appear in the Journal should pertain to proboscideans or to the huge edentate ground-sloths and the aberrant zeuglodons whose bones frequently came to light. Therefore a number of men such as Koch, both Sillimans, J. C. Warren, and others made these forms their chief concern.
Fossil Footprints.—Among the early writers who concerned themselves with these greater fossils was Edward Hitchcock, sometime president of Amherst College, and a geologist of high repute among his contemporaries. Hitchcock is, however, better and more widely known as the pioneer worker on a series of phenomena displayed as in no other place in the region in which he made his home. These are fossil footprints impressed upon the Triassic rocks of the Connecticut valley. It was in the Journal for the year 1836 (29, 307–340) that Hitchcock first called attention to the footmarks, although they had been known and discussed popularly for a number of years previous. James Deane, of Greenfield, was perhaps the first to appreciate the scientific interest of these phenomena, but deeming his own qualifications insufficient properly to describe them, he brought them to the attention of Hitchcock, and the interest of the latter never waned until his death in 1864. Hitchcock wrote paper after paper, publishing many of them in the Journal, again in his Final Report on the Geology of Massachusetts (1841), and later in quarto works, one in the Memoirs of the American Academy of Arts and Sciences and the two others under the authority of the Commonwealth, the Ichnology in 1858, and the Supplement in 1865, the last being a posthumous work edited by his son, Charles H. Hitchcock.
Hitchcock’s conception of the track-makers was more or less imperfect because of the fact that for a long time but a few fragmentary osseous remains were known, either directly or indirectly associated with the tracks, while on the other hand the bird-like character of many of the latter and the discovery of huge flightless birds elsewhere on the globe suggested a very close analogy if not a direct relationship. Hence “bird tracks” they were straightway called, a designation which it has been difficult to remove, even though in 1843 Owen called attention to the need of caution in assuming the existence of so highly organized birds at so early a period, especially when large reptiles were known which might readily form very similar tracks. The footprints are now believed to be very largely of dinosaurian origin, and dinosaurs whose feet corresponded in every detail with the footprints have actually come to light within the same geologic and geographic limitations. This of course refers to the bipedal, functionally three-toed tracks. Of the makers of certain of the obscurer of the quadrupedal trails we are as much in the dark to-day as were the first discoverers of a century ago, so far as demonstrable proof is concerned. We assume, however, that they were the tracks of amphibia and reptiles, beyond which we may not go with certainty.
Agassiz, writing in 1865 (Geological Sketches), says:
“To sum up my opinion respecting these footmarks, I believe that they were made by animals of a prophetic type, belonging to the class of reptiles, and exhibiting many synthetic characters. The more closely we study past creations, the more impressive and significant do the synthetic types, presenting features of the higher classes under the guise of the lower ones, become. They hold the promise of the future. As the opening overture of an opera contains all the musical elements to be therein developed, so this living prelude of the creative work comprises all the organic elements to be successively developed in the course of time.”
Of those whose work was contemporaneous with that of Hitchcock, but one, W. C. Redfield, wrote on Triassic phenomena, and he concerned himself mainly with the fossil fishes of that time, his first paper on this subject appearing in 1837 in the Journal (34, 201), and the last twenty years later.
Paleozoic Vertebrates.—Later the vertebrates of the Paleozoic began to attract attention, footprints from Pennsylvania being described by Isaac Lea, beginning in 1849, a notice of his first paper appearing in the Journal for that year (9, 124). Several papers followed on the reptile Clepsysaurus. Alfred King also wrote on the Carboniferous ichnites, his work slightly antedating that of Lea, but being less authoritative.
But by far the most illuminating of the mid-century writers on Paleozoic vertebrates was Sir William Dawson, a very large proportion of whose numerous papers relate to the Coal Measures of Nova Scotia and their contained plant and animal remains. In 1853 appeared Dawson’s first announcement, written in collaboration with Sir Charles Lyell, of the finding of the bones of vertebrates within the base of an upright fossil tree trunk at South Joggins. These bones were identified by Owen and Wyman as pertaining to a reptilian or amphibian to which the name Dendrerpeton acadianum was given. Following this were several papers published in the Quarterly Journal of the Geological Society, London, describing more vertebrates and associated terrestrial molluscs. In 1863 Dawson summarized his discoveries in the Journal (36, 430–432) under the title of “Air-breathers of the Coal Period,” a paper which was expanded and published under the same title in the Canadian Naturalist and Geologist for the same year. Dawson also printed in the same volume the first account of reptilian(?) footprints from the coal. Thus from time to time there emanated from his prolific pen the account of further discoveries, both in bones and footprints, his final synopsis of the air-breathing animals of the Paleozoic of Canada appearing in 1895. The only other group of vertebrates which claimed his attention were certain whales, on which he occasionally wrote.
Fishes.—The fossil fishes from the Devonian of Ohio found their first exponent in J. S. Newberry, appointed chief geologist of the second geological survey of Ohio, which was established in 1869. These fishes from the Devonian shales belonged for the greater part to the curious group of armored placoderms, the remains of which consist very largely of armor plates with little or no traces of internal skeleton. There was also found in association a shark, Cladoselache, of such marvelous preservation that from some of the Newberry specimens now in the American Museum of Natural History, New York, Bashford Dean has demonstrated the histology of muscle and visceral organs, in addition to the very complete skeletal remains.
Newberry’s work on these forms, begun in 1868, has been carried to further completion by Bashford Dean and his pupil L. Hussakof, as well as by C. R. Eastman. Newberry’s other paleontological work was with the Carboniferous fishes of Ohio, the Carboniferous and Triassic fishes of the region from Sante Fé to the Grand and Green rivers, Colorado, and on the fishes and plants of the Newark system of the Connecticut valley and New Jersey. He also discussed certain mastodon and mammoth remains, and those of the peccary of Ohio, Dicotyles.