The Large Copper (Chrysophanus Dispar)
The brilliant butterfly, figured on Plate 99, is of a coppery orange colour. In the male the fore wings have two black dots in the discal cell, the outer one linear, and the outer margin is narrowly blackish; the hind wings have a linear black mark in the cell, and the outer margin is narrowly edged with blackish and dotted with black. The female is more conspicuously marked with black; there are two, sometimes three, spots in the cell of the fore wings, and a transverse series of seven or eight beyond; the outer margin is broadly bordered with black, and there are generally two spots above the inner angle; the hind wings have a black spot in the cell, and a series of black spots beyond, but the whole basal three-fourths of these wings is often deeply suffused with blackish; the outer margin is bordered and spotted with black. The sexes are much alike on the under side, and have reddish-orange fore wings with bluish grey outer margins, and black spots as on the upper side of the female; the hind wings are bluish-grey, powdered with bluish towards the base, and with whitish ringed black spots; five of these spots are before the linear discal mark, and a series of nine or ten beyond; an orange band on the outer margin has black dots on each edge.
Except as regards the size and the shape of the spots, especially in the female, there appears to have been but little variation noted in this species in England.
The two fine female specimens figured on the plate have a more or less distinct wedge-shaped black spot in the basal end of the discal cell of the fore wings. Dale mentions that he has an “almost entirely black” example of the female in his collection.
The var. rutilus, which is the continental form of our butterfly, is smaller in size, as a rule, the spots are not so large, and the orange band is always narrower on the under side of the hind wings. It has been averred that some of the British specimens are referable to this form.
Newman, writing about 1870, gave the following life-history details:—“The egg is laid on the leaves of the great water-dock (Rumex hydrolapathum) during the month of August, and the young caterpillars (never, to the best of my belief, observed) probably emerge during the following month, and hibernate very early at the base of the petioles.
“The caterpillar is full fed in June, and then lies flat on the dock-leaf, rarely moving from place to place, and, when it does so, gliding with a slug-like motion, the legs and claspers being entirely concealed. The head is extremely small, and can be completely withdrawn into the second segment: the body has the dorsal surface convex, the ventral surface flat; the divisions of the segments are distinctly marked, the posterior margin of each slightly overlapping the anterior margin of the next, and the entire caterpillar having very much the appearance of a Chiton; the sides are slightly dilated, the legs and claspers are seated in closely approximate pairs, nearly on a medio-ventral line. The colour is green, scarcely distinguishable from that of the dock-leaf; there is an obscure medio-dorsal stripe, slightly darker than the disk, and in all probability due to the presence of food in the alimentary canal. The chrysalis is obese, blunt at both extremities, attached by minute hooks at the caudal extremities, and also by a belt round the waist.” Newman adds, “My acquaintance with the caterpillar and chrysalis was made very many years ago in Mr. Doubleday’s garden at Epping, where the very plant of Rumex hydrolapathum, on which the caterpillars fed, is still in existence.”
The caterpillar was described by Stephens, in 1828, as somewhat hairy, bright green, with innumerable white dots. The same author states that the chrysalis was “first green, then pale ash-coloured, with a dark dorsal line and two abbreviated white ones on each side, and, lastly, sometimes deep brown.”
The figure of the caterpillar on Plate 98 is after Westwood, and that of the chrysalis after Newman (“Grammar of Entomology”).
Although he refers to it as “hippothoë,” the Large Copper seems to have been known to Lewin (1795), as he states that specimens had been taken in Huntingdonshire. Haworth (1803) mentions its occurrence in the fens of Cambridgeshire, and Stephens, twenty-five years later, wrote:—“This splendid insect appears to be confined to the fenny counties of Cambridge and Huntingdon, with the neighbouring ones of Suffolk and Norfolk, unless the account of its capture in Wales by Hudson be admitted; but this may probably be the following species [hippothoë], which may, moreover, eventually prove synonymous with Ly. dispar. In the first two localities it appears to occur in great profusion, as several hundred specimens have been captured within these last ten years by the London collectors, who have visited Whittlesea and Yaxley Meres, during the month of July, for the sole purpose of obtaining specimens of this insect.”
Dale states that “the latest capture, consisting of five specimens, appears to have been made in Holme Fen, by Mr. Stretton either in 1847 or 1848.”
There is evidence that floods, which were not uncommon in the home of the Large Copper, were not really injurious to the butterfly, and therefore the occasional submergence of its feeding grounds can hardly have been the cause of its almost sudden destruction. It seems more probable that its disappearance was due to the draining of the fens, and at least it is significant that the two events were almost coincident.
There are records of the butterfly having been taken in various odd localities since it was last seen in fenland, but the latest of those dates back to the year 1865. There seems to be no question that the butterfly is now extinct in England, and, lamentable to relate, the chief locality where we can hope to secure a specimen or two for our collection is in the neighbourhood of Covent Garden, where the only requirement for the capture will be a well-lined purse.
The continental form rutilus is found in Germany, France, Northern Italy, South-Eastern Europe (except Greece), Northern Asia Minor, Armenia, and the Altai. The Asian form auratus occurs in South-Eastern Siberia, Amurland, Corea, Northern China, and Amdo.
Four other kinds of “Coppers” have been reported as occurring in England: these are Chrysophanus hippothoë and C. virgaureæ, both of which have even had English names bestowed upon them, to wit, the Purple-edged Copper and the Scarce Copper; C. gordius, and C. circe (dorilis). These are only mentioned to afford an opportunity for saying that there does not appear to be the least reason for considering either of them to be a British butterfly. Kirby, Barrett, and others, however, think it possible that the first two may have inhabited England in ancient times.